| Ruminants |
Ruminants evolved to consume and subsist on roughage, grasses and shrubs made up largely of cellulose. An understanding of the ruminant digestive system is useful in order to appreciate the dietary requirements of ruminants.
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| The Rumen |
Ruminants have a stomach with four compartments. When a sheep, goat or other ruminant eats, the food initially goes into the first stomach, called the rumen. From there, it moves gradually into the reticulum where it is broken down by bacteria and acid to form cud. Muscles in the reticulum push the cud back into the animals's mouth for a second chewing to help break down the food material even further (rumination). Once the sheep swallows the cud it then enters the omasum for further digestion before it enters the abomasum.
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Diagram showing the four compartments of the
ruminant stomach, viewed from the right hand side |
From the abomasum the food material then enters the small intestine. The abomasum is the ruminant's true, or glandular stomach, and it is very similar to the stomach of non-ruminants, or monogastrics. Combined, these organs occupy almost three quarters of the abdominal cavity, filling most of the left side and extending significantly into the right hand side (Animals with rumen problems often look swollen on the left side due to bloating of the rumen).
When ruminants are born, the rumen is not developed. New-born ruminants only use the abomasum to digest milk, and they are unable to digest plant fibres. As the animals get older and start to eat fibrous foods, the rumen grows and the animals start to ruminate.
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Feed, water and saliva are delivered to the rumen and reticulum (reticulorumen) through the oesophagus. Heavier components in the feed (including foreign objects such as nails and stones) tend to enter the reticulum straight away. Otherwise, the material enters the rumen. The rumen then contains the older fluid-saturated roughage, e.g. from the previous day, towards the bottom, and the newer and lighter feed above this. Fermentation produces a lot of gas and this rises to the top before being belched out through the mouth.
Secretion of fluid into the gastrointestinal tract is of particular importance to herbivorous animals that rely on microbial fermentation for their digestion. A stable environment is important to maintaining a healthy microbial population, and fluid secreted into the gastrointestinal tract, particularly that produced by the salivary gland, is an important contributor to that stability. The saliva provides:
- Fluid for the fermentation vat
- Alkaline buffering. Saliva is rich in bicarbonate, and this buffers the large quantity of acid produced in the rumen and is critical for maintenance of rumen pH.
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| Tables of Requirements |
Below are guidelines for the energy and protein requirements for small livestock in different stages of production. It is unlikely that you will know the actual composition of the diet, especially if there is any element of grazing/browsing/scavenging activity. However, combined with the information in the sections on the nutritive values of different feeds, you may be able to estimate which if any nutrient is limiting in any given situation, and then select the best available feeds for supplying that nutrient.
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| Sheep |
Required concentrations of nutrients (on a dry matter basis)
in the diets of sheep in different states or stages of growth .
Table sourced from:
Carles, A.B. (1983). Sheep production in the tropics. Oxford University Press, Oxford, UK. pp110-111.
Age |
State |
Liveweight |
Daily DM 1 intake |
ME |
Dig. prot 2 |
Ca |
P |
Carotene |
Vitamin D |
|
|
(kg) |
kg |
% liveweight |
(MJ/kg) |
(g/kg) |
(g/kg) |
(g/kg) |
(mg/kg) |
(IU/kg) |
Ewes |
Maintenance |
40 |
0.9 |
2.3 |
8.4 |
48 |
3.2 |
3.0 |
1.8 |
253 |
|
|
50 |
1.0 |
2.0 |
8.4 |
48 |
3.0 |
2.8 |
1.9 |
278 |
|
|
60 |
1.1 |
1.8 |
8.4 |
48 |
2.8 |
2.6 |
2.0 |
303 |
|
|
70 |
1.2 |
1.7 |
8.4 |
48 |
2.7 |
2.5 |
2.2 |
323 |
|
Pregnancy: |
40 |
0.9 |
2.3 |
8.4 |
49 |
2.9 |
2.7 |
1.6 |
245 |
|
weeks 16-21 |
50 |
1.1 |
2.2 |
8.4 |
49 |
2.7 |
2.5 |
1.7 |
253 |
|
and lactation |
60 |
1.3 |
2.1 |
8.4 |
49 |
2.4 |
2.2 |
1.7 |
256 |
|
weeks 9-16 |
70 |
1.4 |
2.0 |
8.4 |
49 |
2.3 |
2.1 |
1.9 |
277 |
|
Lactation: |
40 |
1.5 |
3.7 |
8.8 |
52 |
2.5 |
2.4 |
3.3 |
150 |
|
Singles weeks 1-8 |
50 |
1.7 |
3.3 |
8.8 |
52 |
2.4 |
2.3 |
3.6 |
164 |
|
Twins weeks 9-16 |
60 |
1.9 |
3.2 |
8.8 |
52 |
2.3 |
2.2 |
3.9 |
175 |
|
|
70 |
2.1 |
3.0 |
8.8 |
52 |
2.1 |
2.0 |
4.2 |
185 |
|
Lactation: |
40 |
1.9 |
4.7 |
10.0 |
62 |
5.4 |
3.9 |
2.6 |
117 |
|
Twins weeks 1-8 |
50 |
2.1 |
4.2 |
10.0 |
62 |
5.2 |
3.7 |
3.0 |
132 |
|
|
60 |
2.3 |
3.9 |
10.0 |
62 |
5.0 |
3.6 |
3.3 |
145 |
|
|
70 |
2.5 |
3.6 |
10.0 |
62 |
4.8 |
3.4 |
3.5 |
155 |
|
|
40 |
2.2 |
5.5 |
10.0 |
72 |
5.4 |
3.9 |
2.2 |
100 |
|
|
50 |
2.4 |
4.8 |
10.0 |
72 |
5.2 |
3.7 |
2.6 |
116 |
|
|
60 |
2.6 |
4.3 |
10.0 |
72 |
5.0 |
3.6 |
2.9 |
128 |
|
|
70 |
2.8 |
4.0 |
10.0 |
72 |
4.8 |
3.4 |
3.1 |
139 |
Weaners |
250 |
20 |
1.0 |
5.0 |
10.0 |
67 |
4.6 |
2.6 |
1.3 |
111 |
(ADG3 g) |
180 |
30 |
1.3 |
4.3 |
9.2 |
58 |
4.5 |
2.5 |
1.5 |
128 |
|
120 |
40 |
1.4 |
3.5 |
8.8 |
53 |
4.4 |
2.4 |
1.8 |
159 |
Yearlings |
Diet as for ewes at maintenance, but greater intake, level depending on growth requirement |
1DM: dry matter;
2Dig. prot.: digestible protein;
3ADG: average daily gain.
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Recommended mineral concentrations in the sheep’s diet (on a dry matter basis)
Table sourced from Carles, A.B. (1983). Sheep production in the tropics. Oxford University Press, Oxford, UK. p.112.
Mineral group |
Mineral |
Recommended concentration |
Toxic concentration |
Macromineral |
Sodium |
0.4-1.0 |
|
(g/kg dry matter) |
Calcium |
2.1-5.2 |
|
|
Phosphorus |
1.6-3.7 |
|
|
Magnesium |
0.4-0.8 |
|
|
Potassium |
5.0 |
|
|
Sulphur
|
1.4-2.6 |
|
Micromineral |
Iodine |
0.10-0.80 |
8 |
(mg/kg dry matter) |
Iron |
30-50 |
|
|
Copper |
5 |
8-25 |
|
Molybdenum |
0.5 |
8-20 |
|
Cobalt |
0.1 |
100-200 |
|
Manganese |
20-40 |
|
|
Zinc |
35-50 |
1000 |
|
Selenium |
0.1 |
Over 2 |
|
Fluorine |
|
60-200 or 2 mg/l water |
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| Goats |
Total energy requirements (MJ ME/day) for goats
Data from Peacock, C. (1996). Improving goat production in the tropics. Oxfam/FARM-Africa, Oxford, UK. p.86.
Goat |
Total requirement (maintenance (M) plus growth or activity) |
liveweight (kg) |
M |
Some activity |
Much activity |
Growth |
Pregnancy |
|
|
|
|
50 g/d |
100 g/d |
150 g/d |
|
10 |
2.3 |
2.8 |
3.2 |
4.0 |
5.8 |
7.5 |
5.1 |
15 |
3.2 |
3.8 |
4.4 |
|
|
|
6.9 |
20 |
3.9 |
4.7 |
5.5 |
5.5 |
7.3 |
9.0 |
8.5 |
25 |
4.6 |
5.5 |
6.5 |
|
|
|
10.0 |
30 |
5.3 |
6.4 |
7.4 |
6.8 |
8.6 |
10.3 |
11.5 |
35 |
5.9 |
7.1 |
8.3 |
|
|
|
13.0 |
40 |
6.6 |
7.9 |
9.2 |
8.0 |
9.8 |
11.6 |
14.3 |
45 |
7.2 |
8.6 |
10.1 |
|
|
|
15.6 |
50 |
7.8 |
9.3 |
10.9 |
9.0 |
10.8 |
12.6 |
16.9 |
55 |
8.3 |
10.0 |
11.7 |
|
|
|
18.2 |
60 |
8.9 |
10.7 |
12.5 |
10.3 |
12.0 |
13.8 |
19.4 |
Digestible crude protein (DCP) requirements (g/d) for maintenance and growth of goats
Data from Peacock, C. (1996). Improving goat production in the tropics. Oxfam/FARM-Africa, Oxford, UK. p.86.
Goat |
Total DCP requirement for maintenance (M) and growth |
liveweight |
M |
Growth (g/d) |
Pregnancy |
(kg) |
|
50 |
100 |
150 |
|
10 |
15 |
25 |
35 |
45 |
30 |
20 |
26 |
36 |
46 |
56 |
50 |
30 |
35 |
45 |
55 |
65 |
67 |
40 |
43 |
53 |
63 |
73 |
83 |
50 |
51 |
61 |
71 |
81 |
99 |
60 |
59 |
69 |
79 |
89 |
113 |
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| Poultry |
Simplified nutrient requirements for a laying hen (concentrations in diet on air dry basis)
Table sourced from Smith, A.J. (1990). Poultry. In: Coste, R. (Series editor) The tropical agriculturalist. Macmillan Publishers, London, UK. p.83
Nutrient |
Requirement (range) |
Metabolizable energy (MJ/kg) |
11.5-12.5 |
Crude protein (g/kg) |
165-175 |
Calcium (g/kg) |
25-35 |
Phosphorus (g/kg) |
6-10 |
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Summary of nutrient requirements of chicks, broilers and ducklings (air dry material)
Table sourced from Smith, A.J. (1990). Poultry. In: Coste, R. (Series editor) The tropical agriculturalist. Macmillan Publishers, London, UK. p90.
Constituent |
Bird |
Minimum |
Maximum |
Lysine (g/kg) |
Chicks and ducklings |
10 |
- |
|
Broilers |
11 |
- |
Methionine (g/kg) |
Chicks and ducklings |
4 |
- |
|
Broilers |
4.8 |
- |
Crude fibre (g/kg) |
Chicks and ducklings |
- |
60 |
|
Broilers |
- |
40 |
ME (MJ/kg) |
Chicks |
11.0 |
- |
|
Broilers and ducklings |
13.0 |
- |
Crude protein (g/kg) |
Chicks and ducklings (<4 weeks) |
180 |
- |
|
Ducklings (4-8 weeks) |
160 |
- |
|
Broilers (<5 weeks) |
210 |
- |
|
Broilers (5-10 weeks) |
190 |
- |
Calcium (g/kg) |
|
7 |
13 |
Manganese (mg/kg) |
|
50 |
- |
Sodium chloride (g/kg) |
|
4 |
4 |
Phosphorus (g/kg) |
|
4 |
11 |
Zinc (mg/kg) |
|
40 |
- |
Choline (mg/kg) |
|
1300 |
- |
Niacin (mg/kg) |
|
28 |
- |
Pantothenic acid (mg/kg) |
|
10 |
- |
Riboflavin (mg/kg) |
|
4.0 |
- |
Vitamin A 1 (IU/kg) |
|
1320 |
- |
Vitamin D 3 (IU/kg) |
|
400 |
- |
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Nutrient requirements of laying fowls and laying ducks (air dry material)
Table sourced from Smith, A.J. (1990). Poultry. In: Coste, R. (Series editor) The tropical agriculturalist. Macmillan Publishers, London, UK. p91.
Constituent |
Minimum |
Maximum |
Lysine (g/kg) |
8.0 |
- |
Methionine (g/kg) |
3.5 |
- |
Crude fibre (g/kg) |
- |
70 |
ME (MJ/kg) |
12.0 |
- |
Crude protein (g/kg): |
|
|
Hens |
165 |
- |
Ducks |
190 |
|
Calcium (g/kg) |
30 |
35 |
Manganese (mg/kg) |
30 |
- |
Sodium chloride (g/kg) |
5.0 |
5.0 |
Phosphorus (g/kg) |
5.0 |
10 |
Zinc (mg/kg) |
60 |
- |
Choline (mg/kg) |
1100 |
- |
Niacin (mg/kg) |
8.0 |
- |
Pantothenic acid (mg/kg) |
1.5 |
- |
Riboflavin (mg/kg) |
2.5 |
- |
Vitamin A 1 (IU/kg) |
2700 |
- |
Vitamin D 3 (IU/kg) |
600 |
- |
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Free ranging chickens forage for
natural
sources of nutrients |
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| References and Further Reading |
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| Carles, A.B. (1983). Sheep production in the tropics. Oxford University Press, Oxford |
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| Peacock, C. (1996). Improving goat production in the tropics. Oxfam/FARM-Africa, Oxford |
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| Smith, A.J. (1990). Poultry. In: Coste, R. (Series editor) The tropical agriculturalist. Macmillan Publishers, London |
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